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The Verbenaceae

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Description: Botanical characteristics of this family with photographs and a brief description of a number of individual species.
Flowering Plant Families, UH Botany The Verbenaceae are herbs, shrubs, or trees comprising about 100 genera and 2,600 species that are further characterized by the common occurrence of quadrangular twigs and/or aromatic herbage. The leaves are nearly always opposite or whorled, mostly simple; stipules are lacking. The flowers are nearly always bisexual and zygomorphic. The calyx is synsepalous and most commonly 5-merous. The corolla is sympetalous, usually unequally 5-lobed, and sometimes strongly 2-lipped. The androecium most commonly consists of 4 didynamous stamens adnate to the corolla tube or perigynous zone, alternate with the lobes. The gynoecium consists of a single compound pistil of nearly always 2 carpels, a single terminal or subterminal style, and an unlobed or shallowly lobed ovary, usually with 4 locules (by false septation), each with a single axile ovule. A weakly developed annular nectary disk occurs around the base of the ovary in many species. The fruit is usually a drupe or nutlets.
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